2000 - PhD Molecular Virology, University of London, UK
1985 - HNC Applied Biology, Reading College, UK
1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK
1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
1999 - PhD Botany, University of Otago, NZ
1994 - BSc (Hons) Molecular & Physiological Plant Biology, University of Otago, NZ
2001 - PhD Botany, Department of Plant & Microbial Sciences, University of Canterbury, NZ
1989 - MSc (1st Hons) Botany, Department of Botany, University of Auckland, NZ
1988 - BSc Botany, University of Auckland, NZ
2003 - PhD Neuroscience, University of Regensburg, Germany
1995 - MSc Microbiology, Colorado State University, Colorado, US
1992 - BSc Microbiology, Colorado State University, Colorado, US
1999 - PhD Physiology, University of Alberta, Canada
1984 - BSc Zoology (1st Hons), University of Bristol, UK