1989 - PhD Biotechnology, Durham University, UK
1985 - BSc (Hons) Biotechnology, Leeds University, UK
1999 - PhD Botany, University of Otago, NZ
1994 - BSc (Hons) Molecular & Physiological Plant Biology, University of Otago, NZ
2001 - PhD Botany, Department of Plant & Microbial Sciences, University of Canterbury, NZ
1989 - MSc (1st Hons) Botany, Department of Botany, University of Auckland, NZ
1988 - BSc Botany, University of Auckland, NZ
1999 - PhD Physiology, University of Alberta, Canada
1984 - BSc Zoology (1st Hons), University of Bristol, UK
2002 - PhD Botany, University of Texas, Austin, USA
1991 - B.A. Biology, Swarthmore College, USA
2004 - DSc, Manchester University, UK
1980 - PhD Marine Biology, Manchester University, UK
1976 - BSc (Hons) Marine Biology, Liverpool University, UK
1997 - PhD, Zoology, The University of Reading, UK
1992 - BA (Hons) Natural Sciences, Cambridge University, UK