2000 - PhD Molecular Virology, University of London, UK
1985 - HNC Applied Biology, Reading College, UK
1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK
1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
1976 - MD, Albert Einstein College of Medicine, USA
1973 - AB, Biochemical Sciences, Harvard University, USA
2002 - PhD Molecular Neuroscience, School of Medicine, University of Auckland, NZ
1998 - MSc (1st Hons) Biology and Molecular Genetics, University of Auckland, NZ
1996 - BSc Double Major in Biochemistry and Zoology, University of Auckland, NZ
2000 - PhD Neurobiology & Behavior, SUNY Stony Brook, New York, US
1996 - BSc Psychology, SUNY Stony Brook, New York, US
2006 - Copyediting Certification, UCSD, La Jolla, California, US
1996 - PhD Genetics, MRC Human Genetics Unit and University of Edinburgh, UK
1991 - BSc (Hons) Biotechnology, King's College, University of London, UK