1992 - PhD Biochemistry, University of Kent at Canterbury, UK
1988 - BSc (Hons) Biochemistry, University of Kent at Canterbury, UK
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
1983 - PhD Embryonic Development, Glasgow University, UK
1980 - BA (1st Hons) Zoology, Oxford University, UK
2006 - MRCOphth, Royal College of Ophthalmologists, UK
2002 - MB BChir Medicine, University of Cambridge, UK
2000 - BSc (Hons) Medical Biochemistry, University of Leeds, UK
1984 - PhD Virology , University of Aberdeen, UK
1980 - BSc (Hons) Microbiology, University of Glasgow, UK
1990 - DipLib, Robert Gordon's Institute of Technology, Aberdeen, UK
1992 - PhD Zoology, University of Oxford, UK
1988 - BA (Hons) Zoology, University of Oxford, UK
1996 - MPH Statistics & Epidemiology Modules, University of Dundee, UK
1980 - PhD Pharmacology, University of Edinburgh, UK
1977 - BSc (Biol Sci) (Hons) Pharmacology, University of Edinburgh, UK