1971 - BSc Soil Science, University of Aberdeen, UK
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
2003 - PhD Neurobiology and Physiology, Northwestern University, USA
1998 - MS Molecular Physiology, Vanderbilt University, USA
1996 - BS Biology, Oglethorpe University, USA
1983 - PhD Embryonic Development, Glasgow University, UK
1980 - BA (1st Hons) Zoology, Oxford University, UK
2000 - PhD Developmental Medicine/Biology, Pediatrics, University of Auckland, NZ
1993 - MSc Zoology, University of Auckland, NZ
1989 - BSc Zoology & Cell Biology, University of Auckland, NZ
2005 - PhD Exercise and Sport Science, Tshwane University of Technology, South Africa
1997 - MMedSci Physiology, University of KwaZulu-Natal, South Africa
1994 - BA Hons Human Movement Studies, Rhodes University, South Africa
1997 - PhD Pharmacology, University College London, UK
1990 - MSc Chemical Research, University College London, UK
1989 - BSc (Hons) Applied Chemistry, University of Greenwich, UK