2000 - PhD Molecular Virology, University of London, UK
1985 - HNC Applied Biology, Reading College, UK
1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK
1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK
1996 - PhD Biochemistry, University of Sydney, Australia
1991 - BSc (1st Hons) Biochemistry, University of Sydney, Australia
2006 - PhD Biochemistry/Endocrinology, Queen Mary, University of London, UK
2002 - Post-graduate Certificate “Principles of Protein Structure”, Birkbeck, University of London, UK
2000 - BSc (Hons) Physiology, University of Bristol, UK
2008 - PhD Chemistry, University of Otago, NZ
2002 - BSc (Hons) Chemistry, University of Otago, NZ
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
2000 - PhD Neurobiology & Behavior, SUNY Stony Brook, New York, US
1996 - BSc Psychology, SUNY Stony Brook, New York, US
2006 - Copyediting Certification, UCSD, La Jolla, California, US