1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
2003 - PhD Neurobiology and Physiology, Northwestern University, USA
1998 - MS Molecular Physiology, Vanderbilt University, USA
1996 - BS Biology, Oglethorpe University, USA
1983 - PhD Embryonic Development, Glasgow University, UK
1980 - BA (1st Hons) Zoology, Oxford University, UK
2012 - PhD candidate Developmental Origins of Health and Disease, Massey University, NZ
2004 - Doctor of Veterinary Medicine, Cornell University, USA
1997 - BSc (Hons) Animal Science, Cornell University
1996 - MPH Statistics & Epidemiology Modules, University of Dundee, UK
1980 - PhD Pharmacology, University of Edinburgh, UK
1977 - BSc (Biol Sci) (Hons) Pharmacology, University of Edinburgh, UK
1992 - PhD Zoology, University of Oxford, UK
1988 - BA (Hons) Zoology, University of Oxford, UK
2001 - PhD Neuroscience, McGill University, Canada
1993 - BSc Neurobiology, McGill University, Canada
1990 - Diploma of College Studies, Health Sciences, Vanier College, Montreal, Canada