1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
2003 - PhD Pharmacology, University of Leicester, UK
1999 - BSc (Hons) Pharmacology, University of Manchester, UK
MB ChB (New Zealand Medical Degree), University of Otago, NZ