1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK

1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK

2006 - PhD Biochemistry/Endocrinology, Queen Mary, University of London, UK

2002 - Post-graduate Certificate “Principles of Protein Structure”, Birkbeck, University of London, UK

2000 - BSc (Hons) Physiology, University of Bristol, UK

1970 - PhD Reproductive Biology, University of Liverpool, UK

1967 - MSc Reproduction & Embryology, University of North Wales, UK

1969 - BSc (Hons) Zoology, University of London, UK

2003 - PhD Neurobiology and Physiology, Northwestern University, USA

1998 - MS Molecular Physiology, Vanderbilt University, USA

1996 - BS Biology, Oglethorpe University, USA

1983 - PhD Embryonic Development, Glasgow University, UK

1980 - BA (1st Hons) Zoology, Oxford University, UK

2005 - PhD Exercise and Sport Science, Tshwane University of Technology, South Africa

1997 - MMedSci Physiology, University of KwaZulu-Natal, South Africa

1994 - BA Hons Human Movement Studies, Rhodes University, South Africa

1996 - MPH Statistics & Epidemiology Modules, University of Dundee, UK

1980 - PhD Pharmacology, University of Edinburgh, UK

1977 - BSc (Biol Sci) (Hons) Pharmacology, University of Edinburgh, UK