1970 - PhD Reproductive Biology, University of Liverpool, UK

1967 - MSc Reproduction & Embryology, University of North Wales, UK

1969 - BSc (Hons) Zoology, University of London, UK

1966 - MEng Control Systems, University of Canterbury, NZ

1964 - BScEng Electrical Engineering & Physics, University of Cape Town, South Africa

2003 - PhD Neurobiology and Physiology, Northwestern University, USA

1998 - MS Molecular Physiology, Vanderbilt University, USA

1996 - BS Biology, Oglethorpe University, USA

1983 - PhD Embryonic Development, Glasgow University, UK

1980 - BA (1st Hons) Zoology, Oxford University, UK

1983 - PhD Physics, University of Canterbury, NZ

1980 - MSc Physics, University of Canterbury, NZ

1978 - BSc (Hons) Physics, University of Canterbury, NZ

1997 - PhD Pharmacology, University College London, UK

1990 - MSc Chemical Research, University College London, UK

1989 - BSc (Hons) Applied Chemistry, University of Greenwich, UK

2001 - DSc Chemistry, University of Nottingham, UK

1972 - PhD Chemistry, University of Nottingham, UK

1969 - BSc (Hons) Chemistry, University of Nottingham, UK