2000 - PhD Molecular Virology, University of London, UK

1985 - HNC Applied Biology, Reading College, UK

1992 - PhD Biochemistry, University of Kent at Canterbury, UK

1988 - BSc (Hons) Biochemistry, University of Kent at Canterbury, UK

1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK

1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK

2006 - PhD Biochemistry/Endocrinology, Queen Mary, University of London, UK

2002 - Post-graduate Certificate “Principles of Protein Structure”, Birkbeck, University of London, UK

2000 - BSc (Hons) Physiology, University of Bristol, UK

1970 - PhD Reproductive Biology, University of Liverpool, UK

1967 - MSc Reproduction & Embryology, University of North Wales, UK

1969 - BSc (Hons) Zoology, University of London, UK

2001 - PhD Fisheries Science, Oregon State University, Oregon, US

1995 - MSc (1st Hons) Biological Sciences, Waikato University, NZ

1993 - BSc Biological Sciences (Earth Science minor), Waikato University, NZ

2000 - PhD Neurobiology & Behavior, SUNY Stony Brook, New York, US

1996 - BSc Psychology, SUNY Stony Brook, New York, US

2006 - Copyediting Certification, UCSD, La Jolla, California, US