1992 - PhD Biochemistry, University of Kent at Canterbury, UK

1988 - BSc (Hons) Biochemistry, University of Kent at Canterbury, UK

1970 - PhD Reproductive Biology, University of Liverpool, UK

1967 - MSc Reproduction & Embryology, University of North Wales, UK

1969 - BSc (Hons) Zoology, University of London, UK

2006 - PhD Biochemistry/Endocrinology, Queen Mary, University of London, UK

2002 - Post-graduate Certificate “Principles of Protein Structure”, Birkbeck, University of London, UK

2000 - BSc (Hons) Physiology, University of Bristol, UK

1983 - PhD Embryonic Development, Glasgow University, UK

1980 - BA (1st Hons) Zoology, Oxford University, UK

2005 - PhD Exercise and Sport Science, Tshwane University of Technology, South Africa

1997 - MMedSci Physiology, University of KwaZulu-Natal, South Africa

1994 - BA Hons Human Movement Studies, Rhodes University, South Africa

1999 - PhD Physiology, University of Alberta, Canada

1984 - BSc Zoology (1st Hons), University of Bristol, UK

2012 - PhD candidate Developmental Origins of Health and Disease, Massey University, NZ

2004 - Doctor of Veterinary Medicine, Cornell University, USA

1997 - BSc (Hons) Animal Science, Cornell University