1983 - PhD Embryonic Development, Glasgow University, UK
1980 - BA (1st Hons) Zoology, Oxford University, UK
2000 - PhD Developmental Medicine/Biology, Pediatrics, University of Auckland, NZ
1993 - MSc Zoology, University of Auckland, NZ
1989 - BSc Zoology & Cell Biology, University of Auckland, NZ
1999 - PhD Physiology, University of Alberta, Canada
1984 - BSc Zoology (1st Hons), University of Bristol, UK
2007 - PhD Pharmacology & Toxicology, University of Arkansas for Medical Sciences, US
1997 - BEng Chemical Engineering, Vanderbilt University, US
1996 - MPH Statistics & Epidemiology Modules, University of Dundee, UK
1980 - PhD Pharmacology, University of Edinburgh, UK
1977 - BSc (Biol Sci) (Hons) Pharmacology, University of Edinburgh, UK
2006 - MRCOphth, Royal College of Ophthalmologists, UK
2002 - MB BChir Medicine, University of Cambridge, UK
2000 - BSc (Hons) Medical Biochemistry, University of Leeds, UK
2003 - PhD Microbiology, University of Oxford, UK
1999 - BSc Biological Sciences, University of Leicester, UK