1983 - PhD Embryonic Development, Glasgow University, UK
1980 - BA (1st Hons) Zoology, Oxford University, UK
1996 - MPH Statistics & Epidemiology Modules, University of Dundee, UK
1980 - PhD Pharmacology, University of Edinburgh, UK
1977 - BSc (Biol Sci) (Hons) Pharmacology, University of Edinburgh, UK
1973 - BSc (Hons) French and German, University of Salford, UK
1977 - Member of the Royal Institution of Chartered Surveyors, UK
1987 - BA Philosophy and English Literature, Bucknell University, Lewisburg, PA, USA
MB ChB (New Zealand Medical Degree), University of Otago, NZ
2006 - PhD Molecular Genetics, Neuroscience, University of Cambridge, Downing College, UK
2001 - BSc (1st Hons) Biochemistry & Pharmacology, University of Leeds, UK