1998 - PhD Pharmacology, Monash University, Australia
1993 - BSc (Hons) Pharmacology, Monash University, Australia
1994 - BSC Physiology, University of Auckland, NZ
1971 - BSc Soil Science, University of Aberdeen, UK
1992 - PhD Biochemistry, University of Kent at Canterbury, UK
1988 - BSc (Hons) Biochemistry, University of Kent at Canterbury, UK
1996 - PhD Molecular Genetics, MRC Human Genetics Unit and University of Edinburgh, UK
1992 - MA (Cantab) Natural Sciences, Christ's College, University of Cambridge, UK
1970 - PhD Reproductive Biology, University of Liverpool, UK
1967 - MSc Reproduction & Embryology, University of North Wales, UK
1969 - BSc (Hons) Zoology, University of London, UK
2006 - PhD Biochemistry/Endocrinology, Queen Mary, University of London, UK
2002 - Post-graduate Certificate “Principles of Protein Structure”, Birkbeck, University of London, UK
2000 - BSc (Hons) Physiology, University of Bristol, UK